Definition and Characteristics of Phytohormones:
Phytohormones (plant hormones) are naturally occurring organic chemical messengers synthesized in minuscule concentrations in one part of the plant (such as shoot tips, root tips, or young leaves) and translocated to other parts where they regulate physiological, developmental, and morphogenetic responses.
The Five Major Classes of Plant Hormones:
- Auxins (e.g. Indole-3-Acetic Acid, IAA):
• Synthesized at the shoot apices and young leaves.
• Functions: Promotes cell elongation, initiates root formation in stem cuttings, induces parthenocarpy (seedless fruits in tomatoes), delays leaf abscission, and mediates phototropic and geotropic curvatures.
• Apical Dominance: High auxin concentration in the terminal apical bud suppresses the outgrowth of lateral axillary buds. When the terminal bud is decapitated (e.g. pruning hedges), lateral buds develop into bushy side branches! - Gibberellins (e.g. Gibberellic Acid, GA₃):
• Synthesized in young leaves, developing seeds, and root tips.
• Functions: Promotes phenomenal internodal elongation (reverses genetic dwarfism in pea and corn plants), induces 'bolting' (sudden internode elongation prior to flowering in rosette plants like cabbage), breaks seed and bud dormancy, and stimulates alpha-amylase enzyme secretion during cereal seed germination. - Cytokinins (e.g. Zeatin, Kinetin):
• Synthesized in regions of active cell division (root tips, developing endosperm).
• Functions: Promotes cell division (cytokinesis), overcomes apical dominance, promotes nutrient mobilization, and delays leaf senescence (the Richmond-Lang effect, keeping cut greens fresh longer). - Ethylene ($\text{C}_2\text{H}_4$, Gaseous Hormone):
• Synthesized in ripening fruits and aging leaves.
• Functions: Promotes commercial ripening of fruits (climacteric fruit ripening in bananas, mangoes), promotes transverse swelling of stems, induces female flower formation in cucumbers, and accelerates leaf and fruit abscission. - Abscisic Acid (ABA, 'The Stress Hormone'):
• Synthesized in leaves and stems under drought, cold, or salinity stress.
• Functions: Acts as a growth inhibitor; induces rapid closure of stomata during water stress by stimulating $\text{K}^+$ ion efflux; promotes bud and seed dormancy; accelerates senescence and abscission of leaves and flowers.